Harmony
The Harmony were an intelligent species whom the Hive destroyed.
Overview
The Harmony were a dragon-like species who benefited from the patronage of the Traveler. The Traveler realigned the orbits of ten planets, including the Harmony's homeworld Ana-Harmony, around a black hole, and in the black hole's polar jet it built the Gift Mast, a hollow construct that radiated pure Light and allowed the Harmony to thrive. The Harmonious Flotilla Invincible guarded the Harmony core worlds and the Nicha Thought-ship, a ship of unknown purpose.
Oryx, the Taken King used his newly-built Dreadnaught to destroy the Harmonious Fleet Invincible when the latter gathered to defend the Nicha Thought-ship. Oryx boarded the Nicha Thought-ship to learn the location of the Gift Mast, but was ambushed by the Vex Mind known as Quria, Blade Transform; it is unknown if the Harmony was actively allied with the Vex, or the Vex independently prepared the ambush. After dispatching Quria, Oryx assaulted the Harmony core worlds.
The Harmony weaponized their black hole, using the "Harmony Sting" to fire powerful plasma jets at the Hive fleet. Xivu Arath led the offensive, but her attacks stalled fifty years on as the Harmony turned to "dragon-wishes", allowing their bishops to challenge Xivu Arath directly in her Ascendant Realm, but eventually she was victorious. Savathûn infiltrated Ana-Harmony in secret and set up Hive covens that sowed chaos among the Harmony for a century, and in the process she acquired an unknown secret. Oryx had his broods colonize the rubble of the black hole's accretion disk, allowing them to direct asteroids at the Harmony's fleets and worlds. When they had been weakened sufficiently, he deployed Seeder Ships and invaded the Harmony outright. When Oryx toppled the Gift Mast, the remaining Harmony reportedly despaired and drowned themselves.
Trivia
The description of the Harmony as dragons and their usage of "dragon-wishes" may mean the Harmony were related in some way to the Ahamkara.